Cryo-EM structure of porcine brain ventricles cilia doublet microtubule (48-nm periodicity)
Resolution: 3.65 Å
EM Method: Single-particle
Fitted PDBs: 9cpc
Q-score: 0.378
Leung MR, Sun C, Zeng J, Anderson JR, Niu Q, Huang W, Noteborn WEM, Brown A, Zeev-Ben-Mordehai T, Zhang R
Nature (2025) 637 pp. 1170-1177 [ DOI: doi:10.1038/s41586-024-08337-5 Pubmed: 39743588 ]
- Cilia and flagella associated protein 141 (12 kDa, Protein from Sus scrofa)
- Parkin coregulated (29 kDa, Protein from Sus scrofa)
- Cfap107/c1orf158 (23 kDa, Protein from Sus scrofa)
- Tektin (48 kDa, Protein from Sus scrofa)
- Chromosome 1 c9orf135 homolog (26 kDa, Protein from Sus scrofa)
- Cilia- and flagella-associated protein 53 (62 kDa, Protein from Sus scrofa)
- Ef-hand domain family member b (98 kDa, Protein from Sus scrofa)
- Guanosine-5'-diphosphate (443 Da, Ligand)
- Ciliary microtubule inner protein 2b (28 kDa, Protein from Sus scrofa)
- Outer dynein arm docking complex subunit 3 (72 kDa, Protein from Sus scrofa)
- Ef-hand calcium binding domain 1 (29 kDa, Protein from Sus scrofa)
- Cilia- and flagella-associated protein 52 (68 kDa, Protein from Sus scrofa)
- Tektin (52 kDa, Protein from Sus scrofa)
- Sperm acrosome associated 9 (25 kDa, Protein from Sus scrofa)
- Tubulin beta chain (49 kDa, Protein from Sus scrofa)
- Chromosome 1 c15orf65 homolog (13 kDa, Protein from Sus scrofa)
- Ef-hand domain-containing family member c2 (87 kDa, Protein from Sus scrofa)
- Ciliary microtubule inner protein 2b (30 kDa, Protein from Sus scrofa)
- Piercer of microtubule wall 1 (15 kDa, Protein from Sus scrofa)
- Sperm-associated antigen 8 isoform 1 (50 kDa, Protein from Sus scrofa)
- Cilia and flagella associated protein 77 (32 kDa, Protein from Sus scrofa)
- Cilia-and flagella-associated protein 96 (34 kDa, Protein from Sus scrofa)
- Efcab6 (167 kDa, Protein from Sus scrofa)
- Outer dynein arm-docking complex subunit 4 (78 kDa, Protein from Sus scrofa)
- Enkurin (29 kDa, Protein from Sus scrofa)
- Protein flattop (19 kDa, Protein from Sus scrofa)
- Guanosine-5'-triphosphate (523 Da, Ligand from Sus scrofa)
- Nucleoside diphosphate kinase homolog 7 (42 kDa, Protein from Sus scrofa)
- Tubulin alpha chain (49 kDa, Protein from Sus scrofa)
- Cilia- and flagella-associated protein 20 (22 kDa, Protein from Sus scrofa)
- Coiled-coil domain-containing protein 114 isoform x2 (80 kDa, Protein from Sus scrofa)
- Rib43a domain with coiled-coils 2 (45 kDa, Protein from Sus scrofa)
- Cilia and flagella associated protein 210 (66 kDa, Protein from Sus scrofa)
- Tektin (56 kDa, Protein from Sus scrofa)
- Cilia and flagella associated protein 161 (35 kDa, Protein from Sus scrofa)
- Ef-hand domain containing 1 (74 kDa, Protein from Sus scrofa)
- Tektin (49 kDa, Protein from Sus scrofa)
- Armadillo repeat-containing protein 4 isoform 1 (116 kDa, Protein from Sus scrofa)
- Meiosis-specific nuclear structural protein 1 (60 kDa, Protein from Sus scrofa)
- Cilia from porcine brain ventricles (Cellular component from Sus scrofa)
- Rib43a-like with coiled-coils protein 1 (44 kDa, Protein from Sus scrofa)
- Cilia and flagella associated protein 276 (19 kDa, Protein from Sus scrofa)
- Cilia and flagella associated protein 144 (16 kDa, Protein from Sus scrofa)
- Cilia- and flagella-associated protein 45 (65 kDa, Protein from Sus scrofa)
Neutralization antibody ZCP4C9 bound with SARS-CoV-2 Omicron BA.5 RBD
Resolution: 3.68 Å
EM Method: Single-particle
Fitted PDBs: 8k18
Q-score: 0.153
To Be Published
- Zcp4c9 light chain (11 kDa, Protein from Homo sapiens)
- Spike protein s1 (21 kDa, Protein from Severe acute respiratory syndrome coronavirus 2)
- Zcp4c9 heavy chain (13 kDa, Protein from Homo sapiens)
- Neutralization antibody zcp4c9 bound with sars-cov-2 omicron ba.5 rbd (Complex from Severe acute respiratory syndrome coronavirus 2)
Human spliceosomal PM5 C* complex
Resolution: 2.8 Å
EM Method: Single-particle
Fitted PDBs: 8c6j
Q-score: 0.353
Dybkov O, Preussner M, El Ayoubi L, Feng VY, Harnisch C, Merz K, Leupold P, Yudichev P, Agafonov DE, Will CL, Girard C, Dienemann C, Urlaub H, Kastner B, Heyd F, Luehrmann R
Sci Adv (2023) 9 pp. eadf1785- [ Pubmed: 36867703 DOI: doi:10.1126/sciadv.adf1785 ]
- Peptidyl-prolyl cis-trans isomerase e (33 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein sm d2 (13 kDa, Protein from Homo sapiens)
- Protein bud31 homolog (17 kDa, Protein from Homo sapiens)
- Nitric oxide synthase-interacting protein (33 kDa, Protein from Homo sapiens)
- U5 snrna (36 kDa, RNA from Homo sapiens)
- Pre-mrna-splicing factor slu7 (68 kDa, Protein from Homo sapiens)
- Potassium ion (39 Da, Ligand)
- Splicing factor ess-2 homolog (52 kDa, Protein from Homo sapiens)
- Splicing factor c9orf78 (33 kDa, Protein from Homo sapiens)
- U2 small nuclear ribonucleoprotein b'' (25 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein-associated proteins b and b' (24 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein e (10 kDa, Protein from Homo sapiens)
- Cell division cycle 5-like protein (92 kDa, Protein from Homo sapiens)
- Magnesium ion (24 Da, Ligand)
- Pre-mrna-processing factor 17 (65 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein g (8 kDa, Protein from Homo sapiens)
- Peptidyl-prolyl cis-trans isomerase-like 3 (18 kDa, Protein from Homo sapiens)
- Probable atp-dependent rna helicase ddx41 (69 kDa, Protein from Homo sapiens)
- Pre-mrna-splicing factor syf1 (100 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein sm d1 (13 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein sm d3 (13 kDa, Protein from Homo sapiens)
- Atp-dependent rna helicase dhx8 (139 kDa, Protein from Homo sapiens)
- Pre-mrna-processing factor 19 (55 kDa, Protein from Homo sapiens)
- Human spliceosomal c* complex assembled on pm5 pre-mrna (Complex from Homo sapiens)
- 116 kda u5 small nuclear ribonucleoprotein component (109 kDa, Protein from Homo sapiens)
- Crooked neck-like protein 1 (83 kDa, Protein from Homo sapiens)
- Intron-binding protein aquarius (171 kDa, Protein from Homo sapiens)
- Pleiotropic regulator 1 (57 kDa, Protein from Homo sapiens)
- Pre-mrna-splicing factor syf2 (28 kDa, Protein from Homo sapiens)
- Pre-mrna-splicing factor rbm22 (46 kDa, Protein from Homo sapiens)
- Serine/arginine repetitive matrix protein 2 (300 kDa, Protein from Homo sapiens)
- Peptidyl-prolyl cis-trans isomerase-like 1 (18 kDa, Protein from Homo sapiens)
- U5 small nuclear ribonucleoprotein 40 kda protein (39 kDa, Protein from Homo sapiens)
- Zinc ion (65 Da, Ligand)
- Splicing regulator sde2 (49 kDa, Protein from Homo sapiens)
- Snw domain-containing protein 1 (61 kDa, Protein from Homo sapiens)
- Pre-mrna-processing-splicing factor 8 (273 kDa, Protein from Homo sapiens)
- Pm5 5' exon (78 kDa, RNA from Homo sapiens)
- Nf-kappa-b-activating protein (47 kDa, Protein from Homo sapiens)
- Small nuclear ribonucleoprotein f (9 kDa, Protein from Homo sapiens)
- U5 small nuclear ribonucleoprotein 200 kda helicase (244 kDa, Protein from Homo sapiens)
- Adenosine-5'-triphosphate (507 Da, Ligand)
- U6 snrna (34 kDa, RNA from Homo sapiens)
- Rna-binding protein 8a (19 kDa, Protein from Homo sapiens)
- Spliceosome-associated protein cwc15 homolog (26 kDa, Protein from Homo sapiens)
- Pre-mrna-splicing factor isy1 homolog (33 kDa, Protein from Homo sapiens)
- Inositol hexakisphosphate (660 Da, Ligand)
- Protein fam50a (40 kDa, Protein from Homo sapiens)
- Sting er exit protein (25 kDa, Protein from Homo sapiens)
- Pre-mrna-splicing factor spf27 (26 kDa, Protein from Homo sapiens)
- U2 snrna (60 kDa, RNA from Homo sapiens)
- Prkr-interacting protein 1 (21 kDa, Protein from Homo sapiens)
- U2 small nuclear ribonucleoprotein a' (28 kDa, Protein from Homo sapiens)
- Guanosine-5'-triphosphate (523 Da, Ligand)
- Eukaryotic initiation factor 4a-iii (46 kDa, Protein from Homo sapiens)
- Pre-mrna-splicing factor cwc22 homolog (57 kDa, Protein from Homo sapiens)
- Pm5 intron (74 kDa, RNA from Homo sapiens)
- Splicing factor cactin (88 kDa, Protein from Homo sapiens)
- Protein fam32a (13 kDa, Protein from Homo sapiens)
- Protein mago nashi homolog (17 kDa, Protein from Homo sapiens)
cryo-EM structure of sea urchin sperm doublet microtubule (48-nm periodicity)
Leung MR, Zeng J, Wang X, Roelofs MC, Huang W, Zenezini Chiozzi R, Hevler JF, Heck AJR, Dutcher SK, Brown A, Zhang R, Zeev-Ben-Mordehai T
Cell (2023) 186 pp. 2880- [ Pubmed: 37327785 DOI: doi:10.1016/j.cell.2023.05.026 ]
- Tex33 (40 kDa, Protein from Strongylocentrotus purpuratus)
- Tex43 (14 kDa, Protein from Strongylocentrotus purpuratus)
- Coiled-coil domain-containing protein 105 (49 kDa, Protein from Strongylocentrotus purpuratus)
- Cfap90(c5orf49) (19 kDa, Protein from Strongylocentrotus purpuratus)
- Rib35 (34 kDa, Protein from Strongylocentrotus purpuratus)
- Spata45 (9 kDa, Protein from Strongylocentrotus purpuratus)
- Cilia- and flagella-associated protein 97 (27 kDa, Protein from Strongylocentrotus purpuratus)
- Protein phosphatase 1 regulatory subunit 32 (52 kDa, Protein from Strongylocentrotus purpuratus)
- Flagellar protofilament ribbon protein rib74 (74 kDa, Protein from Strongylocentrotus purpuratus)
- C4orf45 (32 kDa, Protein from Strongylocentrotus purpuratus)
- Tex49 (17 kDa, Protein from Strongylocentrotus purpuratus)
- Cilia- and flagella-associated protein 45 (64 kDa, Protein from Strongylocentrotus purpuratus)
- Nucleoside diphosphate kinase (42 kDa, Protein from Strongylocentrotus purpuratus)
- Spata48 (23 kDa, Protein from Strongylocentrotus purpuratus)
- Outer dense fiber protein 3 (27 kDa, Protein from Strongylocentrotus purpuratus)
- Helo_like_n(loc577943) (34 kDa, Protein from Strongylocentrotus purpuratus)
- Magnesium ion (24 Da, Ligand)
- Cilia- and flagella-associated protein 77 (27 kDa, Protein from Strongylocentrotus purpuratus)
- Guanosine-5'-diphosphate (443 Da, Ligand)
- Cfap276 (19 kDa, Protein from Strongylocentrotus purpuratus)
- Tubulin alpha chain (50 kDa, Protein from Strongylocentrotus purpuratus)
- Flagellar flij protein (14 kDa, Protein from Strongylocentrotus purpuratus)
- Cfap144(fam183a) (16 kDa, Protein from Strongylocentrotus purpuratus)
- Meiosis-specific nuclear structural protein 1 (61 kDa, Protein from Strongylocentrotus purpuratus)
- Meiosis-specific nuclear structural protein 1 (17 kDa, Protein from Strongylocentrotus purpuratus)
- Cilia- and flagella-associated protein 52 (70 kDa, Protein from Strongylocentrotus purpuratus)
- Cilia- and flagella-associated protein 126 (15 kDa, Protein from Strongylocentrotus purpuratus)
- Cfap68(upf0686, c11orf1) (17 kDa, Protein from Strongylocentrotus purpuratus)
- Rib43a-like with coiled-coils protein 2 (45 kDa, Protein from Strongylocentrotus purpuratus)
- Cfa20 domain-containing protein (22 kDa, Protein from Strongylocentrotus purpuratus)
- Outer dense fiber protein 3 (30 kDa, Protein from Strongylocentrotus purpuratus)
- Fam166c (23 kDa, Protein from Strongylocentrotus purpuratus)
- Tektin (49 kDa, Protein from Strongylocentrotus purpuratus)
- C20orf85 (23 kDa, Protein from Strongylocentrotus purpuratus)
- Tex36(cfap95, c9orf135) (25 kDa, Protein from Strongylocentrotus purpuratus)
- Tex26(loc100888047) (19 kDa, Protein from Strongylocentrotus purpuratus)
- Cfap96(c4orf47) (33 kDa, Protein from Strongylocentrotus purpuratus)
- Pacrg (26 kDa, Protein from Strongylocentrotus purpuratus)
- Tektin (46 kDa, Protein from Strongylocentrotus purpuratus)
- Cilia- and flagella-associated protein 161 (32 kDa, Protein from Strongylocentrotus purpuratus)
- Testicular haploid expressed gene protein-like (30 kDa, Protein from Strongylocentrotus purpuratus)
- Coiled-coil domain-containing protein 81 (79 kDa, Protein from Strongylocentrotus purpuratus)
- Atp6v1fnb (21 kDa, Protein from Strongylocentrotus purpuratus)
- Sea urchin sperm doublet microtubule (Cellular component from Strongylocentrotus purpuratus)
- Ef-hand domain-containing family member c2 (85 kDa, Protein from Strongylocentrotus purpuratus)
- Tex49_homologue(loc580808) (20 kDa, Protein from Strongylocentrotus purpuratus)
- Protein fam166b (48 kDa, Protein from Strongylocentrotus purpuratus)
- Pierce1 (13 kDa, Protein from Strongylocentrotus purpuratus)
- Cfap107 (27 kDa, Protein from Strongylocentrotus purpuratus)
- Trichohyalin-plectin-homology domain-containing protein (62 kDa, Protein from Strongylocentrotus purpuratus)
- Saxo3(loc115918676) (37 kDa, Protein from Strongylocentrotus purpuratus)
- Tepp protein (21 kDa, Protein from Strongylocentrotus purpuratus)
- Ef-hand domain-containing protein (66 kDa, Protein from Strongylocentrotus purpuratus)
- Guanosine-5'-triphosphate (523 Da, Ligand)
- Enkurin domain-containing protein (30 kDa, Protein from Strongylocentrotus purpuratus)
- Sperm-associated antigen 8 (23 kDa, Protein from Strongylocentrotus purpuratus)
- Tektin (53 kDa, Protein from Strongylocentrotus purpuratus)
- Trichohyalin-plectin-homology domain-containing protein (65 kDa, Protein from Strongylocentrotus purpuratus)
- Tubulin beta chain (50 kDa, Protein from Strongylocentrotus purpuratus)
Resolution: 3.3 Å
EM Method: Single-particle
Fitted PDBs: 8b0g
Q-score: 0.399
Couves EC, Gardner S, Voisin TB, Bickel JK, Stansfeld PJ, Tate EW, Bubeck D
Nat Commun (2023) 14 pp. 890-890 [ DOI: doi:10.1038/s41467-023-36441-z Pubmed: 36797260 ]
- Complement component c8 beta chain (67 kDa, Protein from Homo sapiens)
- Cd59 glycoprotein (9 kDa, Protein from Homo sapiens)
- Complement component c8 gamma chain (22 kDa, Protein from Homo sapiens)
- Complement component c6 (104 kDa, Protein from Homo sapiens)
- 2c9, cd59 inhibited mac complex (Complex from Homo sapiens)
- Complement component c7 (93 kDa, Protein from Homo sapiens)
- Complement c5 (188 kDa, Protein from Homo sapiens)
- Complement component c8 alpha chain (65 kDa, Protein from Homo sapiens)
- Complement component c9 (63 kDa, Protein from Homo sapiens)
2C9, C5b9-CD59 cryoEM structure
Resolution: 3.3 Å
EM Method: Single-particle
Fitted PDBs: 8b0h
Q-score: 0.405
Couves EC, Gardner S, Voisin TB, Bickel JK, Stansfeld PJ, Tate EW, Bubeck D
Nat Commun (2023) 14 pp. 890-890 [ DOI: doi:10.1038/s41467-023-36441-z Pubmed: 36797260 ]
- Complement component c8 beta chain (67 kDa, Protein from Homo sapiens)
- Complement component c8 gamma chain (22 kDa, Protein from Homo sapiens)
- Complement component c6 (104 kDa, Protein from Homo sapiens)
- 2c9, cd59 inhibited mac complex (Complex)
- Complement component c7 (93 kDa, Protein from Homo sapiens)
- Cd59 glycoprotein (14 kDa, Protein from Homo sapiens)
- Complement component c8 alpha chain (65 kDa, Protein from Homo sapiens)
- Complement c5 (188 kDa, Protein from Homo sapiens)
- Cd59 glycoprotein (Complex from Homo sapiens)
- Complement component c9 (63 kDa, Protein from Homo sapiens)
- Complement components c5, c6, c7, c8 and c9 (Complex from Homo sapiens)
Oligomeric C9 in complex with aE11 Fab
Resolution: 3.2 Å
EM Method: Single-particle
Fitted PDBs: 8de6
Q-score: 0.519
Bayly-Jones C, Ho BHT, Lau C, Leung EWW, D'Andrea L, Lupton CJ, Ekkel SM, Venugopal H, Whisstock JC, Mollnes TE, Spicer BA, Dunstone MA
Commun Biol (2023) 6 pp. 42-42 [ Pubmed: 36639734 DOI: doi:10.1038/s42003-023-04431-y ]
- Ae11 fab vh (15 kDa, Protein from Mus musculus)
- Ae11 fab vl (13 kDa, Protein from Mus musculus)
- Complement component c9 (63 kDa, Protein from Homo sapiens)
- Beta-d-mannopyranose (180 Da, Ligand)
- Quaternary complex of ae11 fab and polyc9 (2 MDa, Complex from Homo sapiens)
- Calcium ion (40 Da, Ligand)
- 2-acetamido-2-deoxy-beta-d-glucopyranose (221 Da, Ligand)
PolyC9 in complex with aE11 Fab
Bayly-Jones C, Ho BHT, Lau C, Leung EWW, D'Andrea L, Lupton CJ, Ekkel SM, Venugopal H, Whisstock JC, Mollnes TE, Spicer BA, Dunstone MA
Commun Biol (2023) 6 pp. 42-42 [ Pubmed: 36639734 DOI: doi:10.1038/s42003-023-04431-y ]
Composite cryo-EM density map of the 48-nm repeat of the human respiratory doublet microtubule
Gui M, Croft JT, Zabeo D, Acharya V, Kollman JM, Burgoyne T, Hoog JL, Brown A
PNAS (2022) 119 pp. e2207605119-e2207605119 [ DOI: doi:10.1073/pnas.2207605119 Pubmed: 36191189 ]
- Cilia- and flagella-associated protein 52 (68 kDa, Protein from Homo sapiens)
- Tubulin alpha-1a chain (50 kDa, Protein from Homo sapiens)
- Uncharacterized protein c15orf65 (13 kDa, Protein from Homo sapiens)
- Protein fam166c (23 kDa, Protein from Homo sapiens)
- Cilia- and flagella-associated protein 20 (22 kDa, Protein from Homo sapiens)
- Protein cfap141 (12 kDa, Protein from Homo sapiens)
- Tektin-1 (48 kDa, Protein from Homo sapiens)
- Protein cfap276 (19 kDa, Protein from Homo sapiens)
- Sperm acrosome-associated protein 9 (25 kDa, Protein from Homo sapiens)
- Enkurin (29 kDa, Protein from Homo sapiens)
- Protein fam183a (16 kDa, Protein from Homo sapiens)
- Magnesium ion (24 Da, Ligand)
- Guanosine-5'-diphosphate (443 Da, Ligand)
- Tektin-3 (56 kDa, Protein from Homo sapiens)
- Parkin coregulated gene protein (29 kDa, Protein from Homo sapiens)
- Nucleoside diphosphate kinase 7 (42 kDa, Protein from Homo sapiens)
- Doublet microtubule and associated proteins (Complex from Homo sapiens)
- Tektin-4 (50 kDa, Protein from Homo sapiens)
- Protein cfap210 (66 kDa, Protein from Homo sapiens)
- Upf0686 protein c11orf1 (17 kDa, Protein from Homo sapiens)
- Tektin-2 (49 kDa, Protein from Homo sapiens)
- Protein cfap95 (26 kDa, Protein from Homo sapiens)
- Protein cfap107 (23 kDa, Protein from Homo sapiens)
- Sperm-associated antigen 8 (51 kDa, Protein from Homo sapiens)
- Uncharacterized protein c5orf49 (17 kDa, Protein from Homo sapiens)
- Meiosis-specific nuclear structural protein 1 (60 kDa, Protein from Homo sapiens)
- Protein flattop (19 kDa, Protein from Homo sapiens)
- Tubulin beta-4b chain (49 kDa, Protein from Homo sapiens)
- Cilia- and flagella-associated protein 53 (61 kDa, Protein from Homo sapiens)
- Ef-hand domain-containing family member b (93 kDa, Protein from Homo sapiens)
- Rib43a-like with coiled-coils protein 2 (45 kDa, Protein from Homo sapiens)
- Guanosine-5'-triphosphate (523 Da, Ligand)
- Protein fam166b (30 kDa, Protein from Homo sapiens)
- Ef-hand domain-containing family member c2 (87 kDa, Protein from Homo sapiens)
- Cilia- and flagella-associated protein 45 (65 kDa, Protein from Homo sapiens)
- Upf0691 protein c9orf116 (15 kDa, Protein from Homo sapiens)
- Cilia- and flagella-associated protein 161 (34 kDa, Protein from Homo sapiens)
- Isoform 2 of cilia- and flagella-associated protein 77 (33 kDa, Protein from Homo sapiens)
- Ef-hand domain-containing protein 1 (74 kDa, Protein from Homo sapiens)
Human coronavirus OC43 spike glycoprotein ectodomain in complex with the 47C9 antibody Fab fragment
Resolution: 3.9 Å
EM Method: Single-particle
Fitted PDBs: 7po5
Q-score: 0.306
Wang C, Hesketh EL, Shamorkina TM, Li W, Franken PJ, Drabek D, van Haperen R, Townend S, van Kuppeveld FJM, Grosveld F, Ranson NA, Snijder J, de Groot RJ, Hurdiss DL, Bosch BJ
Nat Commun (2022) 13 pp. 2921-2921 [ DOI: doi:10.1038/s41467-022-30658-0 Pubmed: 35614127 ]
- Spike glycoprotein (146 kDa, Protein from Human coronavirus OC43)
- 47c9 antibody heavy chain (13 kDa, Protein from Homo sapiens)
- 47c9 antibody light chain (11 kDa, Protein from Homo sapiens)
- 2-acetamido-2-deoxy-beta-d-glucopyranose (221 Da, Ligand)
- Human coronavirus oc43 spike glycoprotein ectodomain in complex with the 47c9 antibody fab fragment (583 kDa, Complex from Human coronavirus OC43)